02802nas a2200289 4500008004100000022001900041245016300060210006900223490000700292520185900299653001202158653001502170653001402185653002502199653002402224653002202248653002502270653001602295653002402311653001602335653001402351100002702365700002002392700002502412700002502437856005002462 2004 eng d aISSN 0342-753600aMultivariate analysis of male genital structures in the Hipparchia semele-muelleri-delattini complex (Nymphalidae, Satyrinae) from the Balkans: how many taxa?0 aMultivariate analysis of male genital structures in the Hipparch0 v273 a

Two closely related Hipparchia taxa, Hipparchia delattini Kudrna, 1975 and H. semele
muelleri Kudrna, 1975 have been described from the Balkans based on differences in male genital
structure, compared to each other and to another nominal European taxon (H. semele Linnaeus, 1758).
Subsequently, Kudrna (1977) synonymised both H. delattini and H. muelleri with H. volgensis
(Mazochin-Porshnjakov, 1952). Application of multivariate statistical techniques on male genital data
indicates a cline in several aspects of genital morphology linking these three taxa across Europe.
Although clusters are repeatedly found that correspond with the three taxa, it is not possible to ascribe
every individual specimen to one of the three Hipparchia taxa. Hipparchia muelleri is shown to occupy
an intermediate position between H. semele and H. delattini. Generally, H. delattini is present in an area
covering part of northern Greece and the central Balkans. H. semele is present in western Europe, the
Balkans and down the western side of Greece. However, individual specimens that classify to H. delattini
in the current analyses may occur much further west, where historically only H. semele has been, and
there appears to be a correlation between putative taxa and altitude with H. delattini occurring at higher
altitudes. It is suggested that genetic differentiation between these taxa has been maintained and enhanced
during glacial-interglacial cycles. The results of this study are discussed in relation to other morphological
characters and biogeography and require further testing with molecular data.

10aBalkans10abiometrics10agenitalia10aHipparchia delattini10aHipparchia muelleri10aHipparchia semele10aHipparchia volgensis10aLepidoptera10anumerical taxonomy.10aNymphalidae10aSatyrinae1 aWakeham-Dawson, Andrew1 aJaksic, Predrag1 aHolloway, Jeremy, D.1 aDennis, Roger, L. H. uhttps://lepidopteragr.myspecies.info/node/714