00397nas a2200109 4500008004100000245007800041210006900119490000800188100001800196700002300214856005000237 1990 eng d00aA survey of the subgenus Plebejides (Sauter, 1986) - preliminary revision0 asurvey of the subgenus Plebejides Sauter 1986 preliminary revisi0 vXII1 aBalint, Zsolt1 aKertesz, Annamaria uhttps://lepidopteragr.myspecies.info/node/71900842nas a2200109 4500008004100000245018800041210007100229490000700300520035700307100001800664856005000682 1993 eng d00aEgy xeromontá n boglárkalepke: a Plebejus pylaon (Fischer von Waldheim, 1832) és rokonsági köre (Lepidoptera, Lycaenidae), III.* A csoport filogenetikája é s állatföldrajza .0 aEgy xeromontá n boglárkalepke a Plebejus pylaon Fischer von Wald0 v373 a

The phylogenetic relationships o f the Plebejides taxa are clarified b y
cladistic methods . The speciation and biogeographic history of the
group is discussed. Notes on the dynamism and the coolescence of the
phyletic branches are given. Detailed English summary and list of
references close the work.

1 aBalint, Zsolt uhttps://lepidopteragr.myspecies.info/node/71800632nas a2200109 4500008004100000245016800041210006900209490000600278520017000284100001800454856005000472 1987 eng d00aDescription of a new subspecies of Plebejus (Plebejides) sephirus Frivaldsky, 1835 from Macedonia with short note on the pylaon-complex (Lepidoptera: Lycaenidae) 0 aDescription of a new subspecies of Plebejus Plebejides sephirus 0 v23 a

Plebejus (Plebejides) sephirus magnificus ssp. n.  is described from Macedonia. The pylaon-complex is shortly discoussed. 
 

1 aBalint, Zsolt uhttps://lepidopteragr.myspecies.info/node/71703997nas a2200145 4500008004100000022001900041245012700060210006900187260001200256490000700268520349000275100001803765700001803783856005003801 1997 eng d aISSN 0722-377300aReformation of the Polyommatus Section with a Taxonomie and Biogeographic OverView (Lepidoptera, Lycaenidae, Polyommatini)0 aReformation of the Polyommatus Section with a Taxonomie and Biog c11.19970 v403 a

The generic composition of the Polyommatus Section of the Polyommatini, originally proposed by Eliot mostly from studies of male genitalia and including 33 valid generic names, is revised.
Recent revisionary works, including study of morphological (and other) characters in both sexes indicates the following genera must be excluded: Cyclargus Nabokov, 1945; Echinargus Nabokov, 1945; Hemiargus Hubner, [1819]; Itylos Draudt, 1921 (= Parachilades Nabokov, 1945); Nabokovia Hemming, 1960 and Pseudochrysops Nabokov, 1945.
Characters are reviewed for the nine genera remaining in Polyommatus Section: Agriades Hubner, [1819]; Albulina Turr, 1909 (= Farsia Zhdanko, 1992, syn. nov.; Pamiria Zhdanko, 1994, syn. nov.; Patricius Balint, 1991, syn. nov. and Plebejidea Koqak, 1993, syn. nov.); Aricia [Reichenbach], 1819 (= Umpria Zhdanko, 1994, syn. nov.);
Chilades Moore, [1881] (= Lachides Nekrutenko, 1984, syn. nov.); Madeleinea Balint, 1993; Paralycaeides Nabokov, 1945; Plebejus Kluk, 1802 (= Alpherakya Zhdanko 1994, syn. nov.); Polyommatus Latreille, 1804 (= Elviria Zhdanko, 1994, syn. nov.) and Pseudolucia Nabokov, 1945.
A list of genera and species is given. Three replacement names are proposed:
Plebejus ardis Balint & Johnson, nom. nov. (replaces Lycaena eversmanni Staudinger, 1894), Plebejus beani Balint & Johnson, nom. nov. (replaces Lycaena indica Evans, 1925) and Plebejus pilgram Balint & Johnson, nom. nov. (replaces Lycaena serica Grum-Grshimailo, 1902).
From examination of type material, the following new species level synonymies are established: Aricia monarchus Higgins, 1981 (junior synonym of Papilio semiargus Rottemburg, 1775), Lycaena idas tshimgana Forster, 1936 (junior synonym of Lycaena argus f. naruena Courvoisier, 1913), Plebejus balinti D’Abrera, 1993 (junior synonym of Lycaena buddhista Alpheraky, 1881), Polyommatus tibetanus Forster, 1940 (junior synonym of Polyommatus annulatus Elwes, 1906) and Vacciniina lornex Higgins, 1981 (junior synonym of Plebejus lucifera selengensis Forster, 1940).
A hypothetical branching diagram of the Polyommatus Section is presented and discussed, with notes concerning historical context (especially regarding the Plebejinae system of Nabokov). The phenomenon of wing discoloration in certain Polyommatini is discussed in light of new data concerning geographic distribution and apparent endemism in various of these taxa. Also examined are historical and current methods of studying the Polyommatini and various views of the tribes’s overall diversity.
In an Appendix, six polyommatine lycaenids are described: Agriades kurtjohnsoni BAum. spec. nov. from Nepal, Albulina gaborronkayi Balint, spec. nov. also from Nepal, Polyommatus frauvartianae spec. nov. from Afghanistan, Polyommatus fabiani spec. nov. from Mongolia, Polyommatus delessei spec. nov. from Iran and Madeleinea gradoslamasi spec. nov. from Peru.

1 aBalint, Zsolt1 aJohnson, Kurt uhttps://lepidopteragr.myspecies.info/node/71302616nas a2200205 4500008004100000245015900041210006900200490000700269520189400276653002002170653002802190653002202218653001902240653002302259653002302282100001602305700002302321700001602344856005002360 2011 eng d00aA combined genetic-morphometric analysis unravels the complex biogeographical history of Polyommatus icarus and Polyommatus celina Common Blue butterflies0 acombined geneticmorphometric analysis unravels the complex bioge0 v203 a

Widespread species have the potential to reveal large-scale biogeographical patterns, as
well as responses to environmental changes possibly unique to habitat generalists. This
study presents a continental-scale phylogeographical analysis of Polyommatus icarus,
one of the most common Palaearctic butterflies, and the morphologically and ecologically
similar Polyommatus celina, a recently discovered cryptic species. By combining data
from mitochondrial [cytochrome c oxidase subunit I (COI)] and nuclear [internal
transcribed spacer (ITS2)] molecular markers with geometric morphometrics, we
document a complex phylogeographical history for the two species. Despite morphological
similarities, the genetic divergence between these two species is high (more than
5% at COI) and they are not sister species. For the first time, we show that P. celina
occurs not only in North Africa but also in Europe, where it inhabits several west
Mediterranean islands, as well as large parts of Iberia, where it occurs in parapatry with
P. icarus. The two species appear to completely exclude each other on islands, but we
provide morphological and molecular evidence that introgression occurred in the Iberian
Peninsula. We discovered strongly diverged lineages that seem to represent relict
populations produced by past range expansions and contractions: Crete and Iberian
isolates for P. icarus, Balearics–Sardinia and Sicily–Lipari for P. celina. This study
shows that a combined genetic-morphometric approach can shed light on cryptic
diversity while providing the necessary resolution to reconstruct a fine-scale phylogeographical
history of species at both spatial and temporal levels.

10acryptic species10ageometric morphometrics10amolecular markers10aPhylogeography10aPolyommatus celina10aPolyommatus icarus1 aDinca, Vlad1 aDapporto, Leonardo1 aVila, Roger uhttps://lepidopteragr.myspecies.info/node/71203386nas a2200289 4500008004100000245013000041210006900171490000700240520249000247653001602737653001702753653001502770653000802785653001702793653002002810653001602830653000902846653001402855653002502869653003402894653002202928653001302950100002802963700002802991700002703019856005003046 2016 eng d00aKaryosystematics and molecular taxonomy of the anomalous blue butterflies (Lepidoptera, Lycaenidae) from the Balkan Peninsula0 aKaryosystematics and molecular taxonomy of the anomalous blue bu0 v103 a

The Balkan Peninsula represents one of the hottest biodiversity spots in Europe. However, the invertebrate
fauna of this region is still insufficiently investigated, even in respect of such well-studied organisms as
Lepidoptera. Here we use a combination of chromosomal, molecular and morphological markers to rearrange
the group of so-called anomalous blue butterflies (also known as ‘brown complex’ of the subgenus
Agrodiaetus Hübner, [1822] and as the Polyommatus (Agrodiaetus) admetus (Esper, 1783) species group)
and to reveal its cryptic taxonomic structure. We demonstrate that P. aroaniensis (Brown, 1976) is not as
widespread in the Balkans as was previously thought. In fact, it has a dot-like distribution range restricted
to the Peloponnese Peninsula in South Greece. Polyommatus orphicus Kolev, 2005 is not as closely related
to the Turkish species P. dantchenkoi (Lukhtanov & Wiemers, 2003) as was supposed earlier. Instead, it
is a Balkan endemic represented by two subspecies: P. orphicus orphicus (Bulgaria) and P. orphicus eleniae
Coutsis & De Prins, 2005 (Northern Greece). Polyommatus ripartii (Freyer, 1830) is represented in the
Balkans by an endemic subspecies P. ripartii pelopi. The traditionally recognized P. admetus (Esper, 1783) is
shown to be a heterogeneous complex and is divided into Polyommatus admetus sensu stricto (the Balkans
and west Turkey) and P. yeranyani (Dantchenko & Lukhtanov, 2005) (east Turkey, Armenia, Azerbaijan
and Iran). Polyommatus nephohiptamenos (Brown & Coutsis, 1978) is confirmed to be a species with a dotlike
distribution range in Northern Greece. Finally, from Central Greece (Timfristos and Parnassos moun-tains) we describe Polyommatus timfristos Lukhtanov, Vishnevskaya & Shapoval, sp. n. which differs by its

haploid chromosome number (n=38) from the closely related and morphologically similar P. aroaniensis
(n=47-48) and P. orphicus (n=41-42). We provide chromosomal evidence for three separate south Balkan
Pleistocene refugia (Peloponnesse, Central Greece and Northern Greece/South Bulgaria) and stress the
biogeographic importance of Central Greece as a place of diversification. Then we argue that the data
obtained have direct implications for butterfly karyology, taxonomy, biogeography and conservation.

10aAgrodiaetus10abiodiversity10achromosome10aCOI10aconservation10acryptic species10aDNA barcode10aITS210akaryotype10amitochondrial marker10aPolyommatus timfristos sp. n.10aprotected species10ared list1 aVishnevskaya, Maria, S.1 aLukhtanov, Vladimir, A.1 aSaifitdinova, Alsu, F. uhttps://lepidopteragr.myspecies.info/node/70401481nas a2200205 4500008004100000245013700041210006900178490000700247520081500254653001601069653001601085653001101101653001601112653001501128653002101143653001701164100002001181700002401201856005001225 2015 eng d00aTaxonomic Position and Status of Polyommatus (Agrodiaetus) iphigenia (Lepidoptera, Lycaenidae) from the Peloponnese, Southern Greece0 aTaxonomic Position and Status of Polyommatus Agrodiaetus iphigen0 v633 a

In our study we use a 690 bp fragment of the COI gene to analyze a taxon from southern
Greece, usually treated as Polyommatus (Agrodiaetus) iphigenia nonacriensis (Brown,
1977). The previous conclusions on taxonomy and nomenclature of P. (A.) iphigenia
nonacriensis were not supported by molecular or cytological data, therefore the problem of
identity of this taxon has remained unsolved. We found that with respect to COI haplotypes,
P. (A.) iphigenia nonacriensis from Greece is similar to the studied populations of P. (A.)
iphigenia (Herrich-Schäffer, 1847) from Turkey and Armenia. Thus, we confirm that the
only Greek Agrodiaetus butterfly with blue wing coloration in males actually belongs to the
species P. (A.) iphigenia.

10aAgrodiaetus10abutterflies10aGreece10aLepidoptera10aLycaenidae10amolecular marker10aPeloponnese.1 aShapoval, Nazar1 aLukhtanov, Vladimir uhttps://lepidopteragr.myspecies.info/node/70303346nas a2200289 4500008004100000245013000041210006900171490000700240520245000247653001602697653001702713653001502730653000802745653001702753653002002770653001602790653000902806653001402815653002502829653003402854653002202888653001302910100002802923700002702951700002802978856005003006 2016 eng d00aKaryosystematics and molecular taxonomy of the anomalous blue butterflies (Lepidoptera, Lycaenidae) from the Balkan Peninsula0 aKaryosystematics and molecular taxonomy of the anomalous blue bu0 v103 a

The Balkan Peninsula represents one of the hottest biodiversity spots in Europe. However, the invertebrate
fauna of this region is still insufficiently investigated, even in respect of such well-studied organisms as
Lepidoptera. Here we use a combination of chromosomal, molecular and morphological markers to rearrange
the group of so-called anomalous blue butterflies (also known as ‘brown complex’ of the subgenus
Agrodiaetus Hübner, [1822] and as the Polyommatus (Agrodiaetus) admetus (Esper, 1783) species group)
and to reveal its cryptic taxonomic structure. We demonstrate that P. aroaniensis (Brown, 1976) is not as
widespread in the Balkans as was previously thought. In fact, it has a dot-like distribution range restricted
to the Peloponnese Peninsula in South Greece. Polyommatus orphicus Kolev, 2005 is not as closely related
to the Turkish species P. dantchenkoi (Lukhtanov & Wiemers, 2003) as was supposed earlier. Instead, it
is a Balkan endemic represented by two subspecies: P. orphicus orphicus (Bulgaria) and P. orphicus eleniae
Coutsis & De Prins, 2005 (Northern Greece). Polyommatus ripartii (Freyer, 1830) is represented in the
Balkans by an endemic subspecies P. ripartii pelopi. The traditionally recognized P. admetus (Esper, 1783) is
shown to be a heterogeneous complex and is divided into Polyommatus admetus sensu stricto (the Balkans
and west Turkey) and P. yeranyani (Dantchenko & Lukhtanov, 2005) (east Turkey, Armenia, Azerbaijan
and Iran). Polyommatus nephohiptamenos (Brown & Coutsis, 1978) is confirmed to be a species with a dotlike
distribution range in Northern Greece. Finally, from Central Greece (Timfristos and Parnassos moun-tains) we describe Polyommatus timfristos Lukhtanov, Vishnevskaya & Shapoval, sp. n. which differs by its haploid chromosome number (n=38) from the closely related and morphologically similar P. aroaniensis (n=47-48) and P. orphicus (n=41-42). We provide chromosomal evidence for three separate south Balkan Pleistocene refugia (Peloponnesse, Central Greece and Northern Greece/South Bulgaria) and stress the biogeographic importance of Central Greece as a place of diversification. Then we argue that the data obtained have direct implications for butterfly karyology, taxonomy, biogeography and conservation.

10aAgrodiaetus10abiodiversity10achromosome10aCOI10aconservation10acryptic species10aDNA barcode10aITS210akaryotype10amitochondrial marker10aPolyommatus timfristos sp. n.10aprotected species10ared list1 aVishnevskaya, Maria, S.1 aSaifitdinova, Alsu, F.1 aLukhtanov, Vladimir, A. uhttps://lepidopteragr.myspecies.info/node/69402553nas a2200145 4500008004100000245015400041210006900195490000700264520199900271100002102270700002802291700002202319700001602341856005002357 2013 eng d00aEstablishing criteria for higher-level classification using molecular data: the systematics of Polyommatus blue butterflies (Lepidoptera, Lycaenidae)0 aEstablishing criteria for higherlevel classification using molec0 v293 a

Most taxonomists agree on the need to adapt current classifications to recognize monophyletic units. However, delineations
between higher taxonomic units can be based on the relative ages of different lineages and ⁄ or the level of morphological
differentiation. In this paper, we address these issues in considering the species-rich Polyommatus section, a group of butterflies
whose taxonomy has been highly controversial. We propose a taxonomy-friendly, flexible temporal scheme for higher-level
classification. Using molecular data from nine markers (6666 bp) for 104 representatives of the Polyommatus section, representing
all but two of the 81 described genera ⁄ subgenera and five outgroups, we obtained a complete and well resolved phylogeny for this
clade. We use this to revise the systematics of the Polyommatus blues, and to define criteria that best accommodate the described
genera within a phylogenetic framework. First, we normalize the concept of section (Polyommatus) and propose the use of subtribe
(Polyommatina) instead. To preserve taxonomic stability and traditionally recognized taxa, we designate an age interval (4–5 Myr)
instead of a fixed minimum age to define genera. The application of these criteria results in the retention of 31 genera of the 81
formally described generic names, and necessitates the description of one new genus (Rueckbeilia gen. nov.). We note that while
classifications should be based on phylogenetic data, applying a rigid universal scheme is rarely feasible. Ideally, taxon age limits
should be applied according to the particularities and pre-existing taxonomy of each group. We demonstrate that the concept of a
morphological gap may be misleading at the genus level and can produce polyphyletic genera, and we propose that recognition of
the existence of cryptic genera may be useful in taxonomy.

1 aTalavera, Gerard1 aLukhtanov, Vladimir, A.1 aPierce, Naomi, E.1 aVila, Roger uhttps://lepidopteragr.myspecies.info/node/69306464nas a2200157 4500008004100000245017900041210006900220490000600289520584000295653003806135653002406173100002206197700001606219700002106235856005006256 2009 eng d00aA comparative analysis of mitochondrial and nuclear DNA sequences in blue butterflies of subgenus Polyommatus (s. str.) Latreille, 1804 (Lepidoptera: Lycaenidae: Polyommatus)0 acomparative analysis of mitochondrial and nuclear DNA sequences 0 v53 a

A molecular phylogenetic analysis of
the genus Polyommatus (s.str.) was carried out using
two independent genetic markers, the mitochondrial
cytochrome c oxidase I (COI) and the nuclear internal
transcribed spacer 2 (ITS2). A 710bp section of COI (5’)
and most of ITS2 (an aligned 638bp) was sequenced for 27
Polyommatus specimens belonging to 17 different taxa. The
Bayesian method was applied to infer phylogenetic trees in
the single gene and combined analyses.
The resulting trees from the single gene analyses
are congruent with each other although their resolution
differs in parts of the tree. The combined analysis produced
separate clusters for most species.
Individuals belonging to Polyommatus icarus
represent one such cluster in the tree. COI- and ITS2
sequences from different subspecies (P. icarus icarus,
P. icarus fuchsia, P. icarus juno) turned out to be similar
throughout its range, from the Iberian Peninsula to
Transbaikalia. We can confirm that the sequences of
P. andronicus are identical to P. icarus. Because both
taxa occur in sympatry, we conclude that the former is a
synonym to the latter: Polyommatus icarus (Rottemburg,
1775) = P. andronicus Coutsis et Ghavalas, 1995 syn. nov.
P. ciloicus and P. icadius form separate clusters in the
tree which confirms that both are separate species from P.
icarus. P. icadius is recorded from Iran for the first time.
The combined molecular phylogenetic analysis of
mitochondrial and nuclear markers helped to clarify the
taxonomy of the P. eros group. It confirmed earlier results
obtained from COI sequences which showed that P. eroides
and P. kamtshadalis are only subspecies of P. eros. ITS2
sequences of P. boisduvalii, P. erotides, P. menelaos and
P. eroides orientalis are identical to P. eros. Therefore we
suggest to rank these taxa as subspecies of P. eros:
– Polyommatus eros boisduvalii (Herrich-Schäffer, 1844):
«Lycaena Boisduvalii» [Herrich-Schäffer, 1844: 121];
«Lycaena ErosVar…Boisduvalii, HS.» [Krulikovsky, 1891:
223 (in Russian)];
«Polyommatus boisduvali (Herrich-Schäffer, 1844)»
[Korshunov, 2002: 161, 385 (in Russian)].
– Polyommatus eros erotides (Staudinger, 1892) stat. rev.:
«Lyc.[aena] Eros O. var. Erotides Stgr.» [Staudinger, 1892:
319];
«Polyommatus … eros… sbsp. erotides Stgr.» [Korshunov,
1972: 363 (in Russian)].
«Polyommatus erotides (Staudinger, 1892)» [Korshunov,
2002: 161, 385 (in Russian)].
– Polyommatus eros menelaos Brown, 1976 stat. nov.:
«Polyommatus menelaos sp. nov.» [Brown, 1976: 82].
Because P. eros eroides is a subspecies of P. eros, the
taxon «orientalis» also becomes a subspecies of P. eros:
– Polyommatus eros orientalis comb. nov.:
«Polyommatus eroides ssp. n. orientalis» [Krzywicki,
1983: 401].
P. forsteri from Iran is very similar to P. eros in
morphology, but these taxa are markedly different in COIand
ITS2 sequences. Therefore we suggest that P. forsteri is
a distinct species.
The taxa P. eros sichuanicus, P. erotides tartarus and
P. tsvetajevi are very similar genetically but form a separate
cluster from P. eros in the single gene and combined analyses.
Therefore we suggest to combine P. eros sichuanicus, P.
erotides tartarus and P. tsvetajevi into a distinct species.
According to our research the valid species name for this
taxon is Polyommatus amorata (Alpheraky, 1897) comb.
nov., stat. nov., a taxon which was originally described as
Lycaena Eros var. Amorata Alph. nova. Its type series is in
the museum of the Zoological Institute in St. Petersburg.
We designate the specimen with the following label as
lectotype: “Sytschuan Mao-Djoov. 26 VIII” is written on
the upperside and “Potanin 1893” on the underside of the
label.
The lectotypes of P. amorata and “P. eros sichuanicus”
are very similar morphologically and these taxa appear to
have identical areas of distribution. Therefore we propose
the following synonymy:
- Polyommatus amorata (Alpheraky, 1897) = P. eros
sichuanicus Murayama, 1983 syn. nov.
P. amorata contains two subspecies which appear to
have some differences in sequences and wing patterns and
are distributed in different areas:
– Polyommatus amorata tartarus (Fruhstorfer, 1916)
comb. nov.:
«Lycaena tithonus tartarus subsp. nov.» [Fruhstorfer,
1916: 19];
«Polyommatus erotides tartarus (Fruhstorfer, 1916)»
[Huang, 1999: 659].
– Polyommatus amorata tsvetajevi (Kurentzov, 1970) stat.
nov.:
«Lycaena tsvetajevi sp. n.» [Kurentzov 1970: 137 (in
Russian)];
«Polyommatus tsvetaevi (Kurentzov, 1970)» [Korshunov,
2002: 161, 385 (in Russian)].
We conclude that the combined analysis of two
independent mitochondrial and nuclear genetic markers
enabled to resolve the taxonomy of a group of closely
related species whose classification had been confused due
to the shortage of available morphological characters.

10aTaxonomy of Polyommatus (s. str.)10aСOI and ITS2 genes1 aVodolazhsky, D.I.1 aWiemers, M.1 aStradomsky, B.V. uhttps://lepidopteragr.myspecies.info/node/69202050nas a2200157 4500008004100000245015800041210006900199490000800268520137900276653011401655100001601769700002101785700002001806700001601826856005001842 2013 eng d00aDispersal, fragmentation, and isolation shape the phylogeography of the European lineages of Polyommatus (Agrodiaetus) ripartii (Lepidoptera: Lycaenidae)0 aDispersal fragmentation and isolation shape the phylogeography o0 v1093 a

Polyommatus ripartii is a biogeographically and taxonomically poorly understood species of butterfly with a
scattered distribution in Europe. Recently, it has been shown that this species includes several European endemic
and localized taxa (galloi, exuberans, agenjoi) that were previously considered species and even protected, a result
that poses further questions about the processes that led to its current distribution. We analysed mitochondrial
DNA and the morphology of P. ripartii specimens to study the phylogeography of European populations. Three
genetically differentiated but apparently synmorphic lineages occur in Europe that could be considered evolutionarily
significant units for conservation. Their strongly fragmented and counterintuitive distribution seems to be the
result of multiple range expansions and contractions along Pleistocene climatic oscillations. Remarkably, based on
the 79 specimens studied, these genetic lineages do not seem to extensively coexist in the distributional mosaic,
a phenomenon most evident in the Iberian Peninsula. One of the important gaps in the European distribution of
P. ripartii is reduced by the discovery of new Croatian populations, which also facilitate a better understanding of
the biogeography of the species.

10abiogeography – conservation – distribution – genetic lineages – morphology – postglacial expansion.1 aDinca, Vlad1 aRunquist, Marten1 aNilsson, Marten1 aVila, Roger uhttps://lepidopteragr.myspecies.info/node/68702176nas a2200109 4500008004100000245017300041210006900214490000800283520162000291653010501911856005002016 2010 eng d00aHow common are dot-like distributions? Taxonomical oversplitting in western European Agrodiaetus (Lepidoptera: Lycaenidae) revealed by chromosomal and molecular markers0 aHow common are dotlike distributions Taxonomical oversplitting i0 v1013 a

Approximately 50 taxa of butterflies in Western Europe have been described as new species or elevated to the level
of species during the last 40 years. Many, especially those belonging to the genus Agrodiaetus, have unusually
localized, ‘dot-like’ distributional ranges. In the present study, we use a combination of chromosomal and molecular
markers to re-evaluate the species status of Agrodiaetus distributed west of the 17th meridian. The results
obtained do not support the current designations of Agrodiaetus galloi, Agrodiaetus exuberans, and Agrodiaetus
agenjoi as endemic species with highly restricted distribution ranges, but indicate that these taxa are more likely
to be local populations of a widely distributed species, Agrodiaetus ripartii. Agrodiaetus violetae is shown to be a
polytypic species consisting of at least two subspecies, including Agrodiaetus violetae subbaeticus comb. nov. and
Agrodiaetus violetae violetae. Agrodiaetus violetae is genetically (but not chromosomally) distinct from Agrodiaetus
fabressei and has a wider distribution in southern Spain than previously believed. Agrodiaetus humedasae from
northern Italy is supported as a highly localized species that is distinct from its nearest relatives. We propose a
revision of the species lists for Agrodiaetus taking these new data into account. The results reported in the present
study are relevant to animal conservation efforts in Europe because of their implications for IUCN Red List
priorities.

10asystematics – species – conservation – distribution – range – reinforcement – phylogeny. uhttps://lepidopteragr.myspecies.info/node/68501004nas a2200097 4500008004100000245018500041210007100226490000700297520055200304856005000856 1993 eng d00aEgy xeromontán boglárkalepke: a Plebejus pylaon (Fischer von Waldheim, 1832) és rokonsági köre (Lepidoptera, Lycaenidae), III.* A csoport filogenetikája és állatföldrajza.0 aEgy xeromontán boglárkalepke a Plebejus pylaon Fischer von Waldh0 v373 a

BÁLINT, Zsolt: A xeromontane lycaenid butterfly species: Plebejus
plyaon (Fischer von Waldheim, 1832) and its relatives. Part III: The
phylogeny and biogeography of the group.
The phylogenetic relationships of the Plebejides taxa are clarified by
cladistic methods. The speciation and biogeographic history of the
group is discussed. Notes on the dynamism and the coolescence of the
phyletic branches are given. Detailed English summary and list of
references close the work.

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