02802nas a2200289 4500008004100000022001900041245016300060210006900223490000700292520185900299653001202158653001502170653001402185653002502199653002402224653002202248653002502270653001602295653002402311653001602335653001402351100002702365700002002392700002502412700002502437856005002462 2004 eng d aISSN 0342-753600aMultivariate analysis of male genital structures in the Hipparchia semele-muelleri-delattini complex (Nymphalidae, Satyrinae) from the Balkans: how many taxa?0 aMultivariate analysis of male genital structures in the Hipparch0 v273 a

Two closely related Hipparchia taxa, Hipparchia delattini Kudrna, 1975 and H. semele
muelleri Kudrna, 1975 have been described from the Balkans based on differences in male genital
structure, compared to each other and to another nominal European taxon (H. semele Linnaeus, 1758).
Subsequently, Kudrna (1977) synonymised both H. delattini and H. muelleri with H. volgensis
(Mazochin-Porshnjakov, 1952). Application of multivariate statistical techniques on male genital data
indicates a cline in several aspects of genital morphology linking these three taxa across Europe.
Although clusters are repeatedly found that correspond with the three taxa, it is not possible to ascribe
every individual specimen to one of the three Hipparchia taxa. Hipparchia muelleri is shown to occupy
an intermediate position between H. semele and H. delattini. Generally, H. delattini is present in an area
covering part of northern Greece and the central Balkans. H. semele is present in western Europe, the
Balkans and down the western side of Greece. However, individual specimens that classify to H. delattini
in the current analyses may occur much further west, where historically only H. semele has been, and
there appears to be a correlation between putative taxa and altitude with H. delattini occurring at higher
altitudes. It is suggested that genetic differentiation between these taxa has been maintained and enhanced
during glacial-interglacial cycles. The results of this study are discussed in relation to other morphological
characters and biogeography and require further testing with molecular data.

10aBalkans10abiometrics10agenitalia10aHipparchia delattini10aHipparchia muelleri10aHipparchia semele10aHipparchia volgensis10aLepidoptera10anumerical taxonomy.10aNymphalidae10aSatyrinae1 aWakeham-Dawson, Andrew1 aJaksic, Predrag1 aHolloway, Jeremy, D.1 aDennis, Roger, L. H. uhttps://lepidopteragr.myspecies.info/node/71403740nas a2200313 4500008004100000245011000041210006900151260001700220490000700237520281700244653001703061653000803078653002603086653001203112653000903124653001603133653002003149653002303169653001803192653002003210100003003230700001703260700001703277700002103294700002103315700002403336700001603360856005003376 2011 eng d00aTracing the origin of disjunct distributions: a case of biogeographical convergence in Pyrgus butterflies0 aTracing the origin of disjunct distributions a case of biogeogra cOctober 20110 v383 a

Aim To study the biogeographical factors responsible for the current disjunct
distributions of two closely related species of butterflies (Pyrgus cinarae and
Pyrgus sidae, Lepidoptera: Hesperioidea). Both species have small populations in
the Iberian Peninsula that are isolated by more than 1000 km from their nearest
conspecifics. Because these species possess similar ecological preferences and
geographical distributions, they are excellent candidates for congruent
biogeographical histories.
Location The Palaearctic region, with a special focus on the Mediterranean
peninsulas as glacial refugia.
Methods We integrated phylogeography and population genetic analyses with
ecological niche modelling. The mitochondrial gene cytochrome c oxidase
subunit 1 (COI) and the non-coding nuclear marker internal transcribed spacer 2
(ITS2) were analysed for 62 specimens of P. cinarae and for 80 of P. sidae to infer
phylogeography and to date the origin of disjunct distributions. Current and
ancestral [Last Glacial Maximum using MIROC (Model for Interdisciplinary
Research on Climate) and CCSM (Community Climate System Model)
circulation models] distribution models were calculated with Maxent. Using
present climatic conditions, we delimited the ecological space for each species.
Results The genetic structure and potential ancestral distribution of the two
species were markedly different. While the Iberian population of P. cinarae had an
old origin (c. 1 Ma), that of P. sidae was closely related to French and Italian
lineages (which jointly diverged from eastern populations c. 0.27 Ma). Ecological
niche modelling showed that minor differences in the ecological preferences of
the two species seem to account for their drastically different distributional
response to the last glacial to post-glacial environmental conditions. Although the
potential distribution of P. cinarae was largely unaffected by climate change,
suitable habitat for P. sidae strongly shifted in both elevation and latitude. This
result might explain the early origin of the disjunct distribution of P. cinarae, in
contrast to the more recent disjunction of P. sidae.
Main conclusions We show that convergent biogeographical patterns can be
analysed with a combination of genetic and ecological niche modelling data. The
results demonstrate that species with similar distributional patterns and ecology
may still have different biogeographical histories, highlighting the importance of
including the temporal dimension when studying biogeographical patterns.

10aBiogeography10aCOI10adisjunct distribution10aecology10aITS210aLepidoptera10aniche modelling10aPalaearctic region10apalaeoclimate10aphylogeography.1 aHernandez-Roldan, Juan, L1 aMurria, Cesc1 aRomo, Helena1 aTalavera, Gerard1 aZakharov, Evgeny1 aHebert, Paul, D. N.1 aVila, Roger uhttps://lepidopteragr.myspecies.info/node/70501481nas a2200205 4500008004100000245013700041210006900178490000700247520081500254653001601069653001601085653001101101653001601112653001501128653002101143653001701164100002001181700002401201856005001225 2015 eng d00aTaxonomic Position and Status of Polyommatus (Agrodiaetus) iphigenia (Lepidoptera, Lycaenidae) from the Peloponnese, Southern Greece0 aTaxonomic Position and Status of Polyommatus Agrodiaetus iphigen0 v633 a

In our study we use a 690 bp fragment of the COI gene to analyze a taxon from southern
Greece, usually treated as Polyommatus (Agrodiaetus) iphigenia nonacriensis (Brown,
1977). The previous conclusions on taxonomy and nomenclature of P. (A.) iphigenia
nonacriensis were not supported by molecular or cytological data, therefore the problem of
identity of this taxon has remained unsolved. We found that with respect to COI haplotypes,
P. (A.) iphigenia nonacriensis from Greece is similar to the studied populations of P. (A.)
iphigenia (Herrich-Schäffer, 1847) from Turkey and Armenia. Thus, we confirm that the
only Greek Agrodiaetus butterfly with blue wing coloration in males actually belongs to the
species P. (A.) iphigenia.

10aAgrodiaetus10abutterflies10aGreece10aLepidoptera10aLycaenidae10amolecular marker10aPeloponnese.1 aShapoval, Nazar1 aLukhtanov, Vladimir uhttps://lepidopteragr.myspecies.info/node/70302409nas a2200169 4500008004100000245011100041210006900152490000800221520184500229653002502074653001602099653001502115653001902130100002102149700001902170856005002189 2007 eng d00aThe effects of Pleistocene glaciations on the phylogeography of Melitaea cinxia (Lepidoptera: Nymphalidae)0 aeffects of Pleistocene glaciations on the phylogeography of Meli0 v1043 a

Partial (600 bp) sequences of mitochondrial cytochrome oxidase I (COI) gene were used to infer the phylogeography of
Melitaea cinxia (Lepidoptera: Nymphalidae) across the entire distributional range of the species, encompassing north Africa and
Eurasia. Cladistic analysis of 49 distinct haplotypes (haplotype and nucleotide diversity were 0.95 and 0.027, respectively) revealed
strong phylogeographic structure in M. cinxia, characterised by four major clades: Morocco; Western (Iberia, France, Italy); Central
(central and northern Western Europe, Balkans, Greece, Anatolia, Levant); and Eastern (eastern Baltic, Urals, Iran, Siberia, China);
separated by average pairwise distances of beween 2 and 6 percent. This pattern is consistent with the location of southern glacial
refugia in the Iberian, Italian and Balkan peninsulas, as well as multiple eastern refugia. The Western clade is further structured into
south-central Iberian, northern Iberian (and French) and southern Italian sub-clades; and the Eastern clade into Near Eastern and Far
Eastern sub-clades; with weaker phylogeographical concordance within the Central clade, except for a large area in central and
northern Western Europe which is monomorphic for COI haplotype. The Baltic and eastern Europe have been primarily colonized
by the Far Eastern sub-clade, rather than the Central (Balkan) clade, highlighting the importance of including Near and Far Eastern
populations in phylogeographic studies of Palearctic species. Maps showing the extent of clades and sub-clades suggest several
regions of secondary contact and possible hybridization. Interspecific comparison of representative M. cinxia haplotypes supports a
monophyletic origin of all M. cinxia.

10acytochrome oxidase I10aLepidoptera10aPalearctic10aPhylogeography1 aWahlberg, Niklas1 aSaccheri, Ilik uhttps://lepidopteragr.myspecies.info/node/70201796nas a2200193 4500008004100000245015800041210006900199490000700268520114200275653001401417653002101431653001601452653001701468653001601485653001801501100001701519700001601536856005001552 2001 eng d00aButterfly species-group taxa from the Balkans and western Anatolia attributed to Imre Frivaldszky (1799-1870) (Lepidoptera: Hesperioidea & Papilionoidea)0 aButterfly speciesgroup taxa from the Balkans and western Anatoli0 v933 a

Twenty-one butterfly species group taxa, the authorship of which has been traditionally attributed to FRIVALDSZKY by classical XlXth century lepidopterists, though validly described for the first time either by FREYER (eleven taxa), HERRICII-SCHÄFFER (seven taxa) or FRIVALDSZKY himself (three taxa), are discussed in the context of historical material provided to the first two workers, either directly or indirectly, by FRIVALDSZKY and referred to as such in their works as well as in those of other contemporary authors. Special attention is further paid to the Frivaldszky Insect Collection, currently deposited in the Hungarian Natural History Museum and to its Catalogue (FRIVALDSZKY 1864a). Five nomina nuda have been published by FRIVALDSZKY before any valid and available description appeared for the taxon in question, eight afterwards. Syntypes are determined here for the nominal taxa Lycena [sic!] eroides FRIVALDSZKY, 1835 and Chilades trochylus var. grisea ABAFI-AlGNER, 1906, while the type locality of nine nominal taxa is either determined or corrected and one new synonym is established. With 15 figures.

10aAnatolia.10aBalkan Peninsula10aFrivaldszky10aHesperioidea10aLepidoptera10aPapilionoidea1 aÎ’alint, Zs.1 aOlivier, A. uhttps://lepidopteragr.myspecies.info/node/681